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Darwin Correspondence Project

From J. T. Riches   12 November 1877

4 Avon Terrace | Defoe Road | Lower Tooting | S.W.

12.11.77.

Sir.

I think you do not mention the genus Comparettia in your excellent work on Fertilization in Orchids etc. I have just had the opportunity of examining the structure of C. falcata, and find that it is frequently self-fertilizing1   Out of 5 flowers, there were not less than 4 with the ovaries well fertilised. and in the other flower, I found the pollen masses upon the stigma. It seems that when the stigma is ready the anther ruptures from the base upwards, and by the least shake or disturbance the heavy pollen masses are dislodged, and, the rostellum being very narrow and beak-like, there is nothing to prevent it from falling upon the stigma. the caudicles, not being able to sustain their weight,; the flexibility of the latter I consider is very important to effect cross-fertilization; but, that, cross-fertilisation may be brought about the pollen mass must be removed before the complete rupture of the anther has taken place. The naked disk at the end of the rostellum is very viscid, and firmly attaches itself to any intruder in about 30 seconds; so firm in fact, that I could not blow it of. if the caudicle remained such when drawn, it would not place the pollen against the stigma, but owing to their bending the stigma is reached.2

I should be glad if you would correct any errors I may have made.. and you are at perfect liberty to copy the rough sketches, and, if you will kindly return. the originals,3 as that is what I may require, you will oblige | faithfully | your’s | J. T. Riches

C. Darwin Esqr.

Footnotes

Comparettia falcata (the snail orchid or sickle-leaved comparettia) is an epiphytic orchid found in Mexico, Central America, the West Indies, and South America.
Riches uses CD’s terminology to describe the relevant floral organs (for CD’s explanation of his usage, see Orchids 2d ed., pp. 3–5).
No sketches or copies of sketches were found with this letter.

Bibliography

Orchids 2d ed.: The various contrivances by which orchids are fertilised by insects. By Charles Darwin. 2d edition, revised. London: John Murray. 1877.

Summary

Has studied the Comparettia falcata, not mentioned in Orchids, and found it is often self-fertilising.

Letter details

Letter no.
DCP-LETT-11231
From
John Thomas Riches
To
Charles Robert Darwin
Sent from
Lower Tooting
Source of text
DAR 176: 154
Physical description
ALS 3pp

Please cite as

Darwin Correspondence Project, “Letter no. 11231,” accessed on 19 April 2024, https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-11231.xml

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