To J. D. Hooker 11 March [1858]
Down Bromley Kent
March 11th
My dear Hooker
I have been thinking over your note of this morning, which like all your others is of real value to me.— I quite agree with observation on small local genera not varying & mundane ones varying; but this instead of an objection strikes me as rather the very point, which I want to show by my tables.
But I will not discuss the whole subject here; indeed I am trying,as far as able, not to settle my conviction till all my results are finally worked out; when, if you will let me, I will have my M.S. fairly copied out, & sent to you.—
I really cannot at present doubt that for my theoretical object that local Floras are the best; my object being to see whether varieties are not incipient species.—1 Perhaps you will see my point of view, by taking a strong case; viz Marsupialia might well be an increasing group in Australia & a diminishing one when put into competition with Placentalia in S. America; if I lumped them, the results would be confounded & false for both sides.—
Of course I do not suppose with groups of plants so widely extended as they are, that there ever shd. be such difference, as there might be in case of Mammals. Therefore I agree that orders in a Prodromus not obeying my rule as with Labiatæ & Verbenaceæ is a serious objection; though not nearly so fatal, in my opinion, if in a local Flora.—2 I was led to all this work by a remark of Fries, that the species in large genera, were more closely related to each other than in small genera;3 & I thought if this were so, seeing that varieties & species are so hardly distinguishable, I concluded that I shd. find more varieties in the large genera than in the small: but at first, seeing the many causes of doubt, I certainly did not expect to find more than three-fourths of the Floras, yielding the result, which they have.— But I will not go on; as someday I hope you will read my short discussion on whole subject.—
You have done me infinite service, whatever opinion I come to, in drawing my attention to at least the possibility or the probability of Botanists recording more varieties in the large than in the small genera.—4 It will be hard work for me to be candid in coming to my conclusion.—
Ever yours most truly | C. Darwin
I shall be several weeks at my present job.5
The work has been turning out badly for me this morning & I am sick at heart & oh my God how I do hate species & varieties.
P.S.6 I see that I had not understood your figures— They are quite sufficent, (so I do not care for Book) to show how hostile the result is—7 But I look at this as too small an order as any good test; I find to make Balance such as I have always later, there wd be only 4 genera on one side & one of these gigantic; I very well know how you will sneer at this wriggling out— C. D.
I shall of course allude to your result on Urticeæ8
Footnotes
Bibliography
Candolle, Augustin Pyramus de and Candolle, Alphonse de. 1824–73. Prodromus systematis naturalis regni vegetabilis, sive enumeratio contracta ordinum generum specierumque plantarum huc usque cognitarum, juxta methodi naturalis normas digesta. 19 vols. Paris: Treuttel & Würtz [and others].
Fries, Elias Magnus. 1850. A monograph of the Hieracia; being an abstract of Prof. Fries’s ""Symbolæ ad Historiam Hieraciorum"". Botanical Gazette 2: 85–92, 185–8, 203–19. [Vols. 5,7]
Natural selection: Charles Darwin’s Natural selection: being the second part of his big species book written from 1856 to 1858. Edited by R. C. Stauffer. Cambridge: Cambridge University Press. 1975.
Weddell, Hugh Algernon. 1856. Monographie de la famille des Urticés. Paris.
Summary
JDH’s "objection" that small local genera do not vary and mundane ones do, is exactly CD’s point. Local floras useful to test idea that varieties are incipient species. Same genus in different countries cannot be lumped.
Letter details
- Letter no.
- DCP-LETT-2239
- From
- Charles Robert Darwin
- To
- Joseph Dalton Hooker
- Sent from
- Down
- Source of text
- DAR 114: 228
- Physical description
- ALS 9pp
Please cite as
Darwin Correspondence Project, “Letter no. 2239,” accessed on 26 September 2022, https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-2239.xml
Also published in The Correspondence of Charles Darwin, vol. 7